Altitudinal zonation in mountainous regions describes the natural layering of ecosystems that occurs at distinct altitudes due to varying environmental conditions. Temperature, humidity, soil composition, and solar radiation are important factors in determining altitudinal zones, which consequently support different vegetation and animal species. Altitudinal zonation was first hypothesized by geographer Alexander von Humboldt who noticed that temperature drops with increasing elevation. Zonation also occurs in intertidal and marine environments, as well as on shorelines and in wetlands. Scientist C.